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Gordon Warren, PhD

Distinguished University Professor    

Ph.D.,Exercise Physiology
University of Georgia, Athens, Georgia (1989)

M.S., Biomedical Engineering
Massachusetts Institute of Technology, Cambridge, Massachusetts (1986)

B.S., Nuclear Engineering
Georgia Institute of Technology, Atlanta, Georgia (1980)


Office Location

1281 Urban Life

Courses Taught

  • Research I: Methods
  • Research II: Investigations
  • Research III: Synthesis
  • Research IV: Dissemination
  • Systematic Review and Meta-Analysis
  • Exercise Physiology
  • Pathophysiology

Selected Publications

Willett, N.J., L. Krishnan, M.T. Li, R.E. Guldberg, and G.L. Warren. Guidelines for models of skeletal muscle injury and therapeutic assessment. Cells Tissues Organs. In press.

Nichenko, A.S., W.M. Southern, M. Atuan, J. Luan, K.B. Peissig, S.J. Folz, A.M. Beedle, G.L. Warren, and J.A. Call. Mitochondrial maintenance via autophagy contributes to functional skeletal muscle regeneration and remodeling. Am. J. Physiol: Cell Physiol. Epub ahead of print. DOI: 10.1152/ajpcell.00066.2016.

Warren, G.L., J.A. Call, A.K. Farthing, and B. Baadom-Piaro. Minimal evidence for a secondary loss of strength after an acute muscle injury: A systematic review and meta-analysis. Sports Med. Epub ahead of print. DOI: 10.1007/s40279-016-0528-7.

Collier, N.B., M.A. Hardy, M.L. Millard-Stafford, and G.L. Warren. Small beneficial effect of caffeinated energy drink ingestion on strength. J. Strength Cond. Res. 30: 1862-1870, 2016.

Henes, S.T., A. Johnson, M. Toner, K. Mamaril, M. Kelkar, Y. Xiao, and G.L. Warren. Assessing resting metabolic rate in overweight and obese adolescents with a portable indirect calorimeter: A pilot study for validity and reliability. Nutr. Clin. Pract. 31: 355-361, 2016.

Morelli, K.M., J.D. Garcia, L.B. Brown, A.K. Farthing, J.L. Miller, and G.L. Warren. Clinical measures predicting knee extensor muscle activation during a maximal voluntary isometric contraction. J. Exerc. Sports Orthop. 3: 1-10, 2016.

Morelli, K.M., M. Carrelli, M.A. Nunez, C. Smith, and G.L. Warren. Addition of hip exercises to treatment of patellofemoral pain syndrome: A meta-analysis. World J. Meta-Anal. 3: 118-124, 2015.

Novotny, S.A., G.L. Warren, and M.W. Hamrick. Aging and the muscle-bone relationship. Physiology. 30: 8-16, 2015.

Mader, T.L., S.A. Novotny, A.S. Lin, R.E. Guldberg, D.A. Lowe, and G.L. Warren. CCR2 elimination in mice results in larger and stronger tibial bones but bone loss is not attenuated after ovariectomy or denervation. Calc. Tissue Int. 95: 457-466, 2014.

Novotny, S.A., T.L. Mader, A.G. Greising, A.S. Lin, R.E. Guldberg, G.L. Warren, and D.A. Lowe. Low intensity, high frequency vibration training to improve musculoskeletal function in a mouse model of Duchenne muscular dystrophy. PloS One. 9:e104339. doi: 10.1371/journal.pone.0104339, 2014.

Chen, Y.P., S. Pope, D. Tyler, and G.L. Warren. Effectiveness of constraint-induced movement therapy on upper-extremity function in children with cerebral palsy: A systematic review and meta-analysis of randomized controlled trials. Clin. Rehab. 10: 938-953, 2014.

Li, M.T., N.J. Willett, B.A. Uhrig, R.E. Guldberg, and G.L. Warren. Functional analysis of limb recovery following autograft treatment of volumetric muscle loss in the quadriceps femoris. J. Biomech. 47: 2013-2021, 2014.

Call, J.A., G.L. Warren, M. Verma, and D.A. Lowe. Acute failure of action potential conduction in mdx muscle reveals new mechanism of contraction-induced force loss. J. Physiol. 591: 3765-3776, 2013.

Willett, N.J.,M.A. Li, B.A. Uhrig, J.D. Boerckel, G.L. Warren, and R.E. Guldberg. Attenuated BMP-2 mediated bone regeneration in a rat model of composite bone and muscle injury. Tissue Eng. Part C Methods. 19: 316-325, 2013.


  • Work-/exercise-induced and traumatic injuries of skeletal muscle
  • Functional interactions of bone and skeletal  muscle in healthy, diseased, and injured conditions
  • Estrogen’s effect on skeletal muscle function
  • Factors affecting skeletal muscle activation (e.g., caffeine, body posture, clinical measures)


“Combining Clinically Available Rehabilitative and Regenerative Strategies to Improve Functional Outcomes after Severe Extremity Trauma” DOD grant in collaboration with the U.S. Army Institute of Surgical Research, 2016-2019.

“Interaction of Estrogen, Age and Activity on Musculoskeletal Strength in Females” NIH R01 grant in collaboration with the University of Minnesota, 2009-2019.

Professional Service Activities

American Federation for Aging Research (AFAR) National Scientific Advisory Council, Committee member, 2005-present.

Served as manuscript reviewer for the American Journal of Physiology, Archives of Physical Medicine and Rehabilitation, Aviation, Space, & Environmental Med Biological Research for Nursing, Canadian Journal of Applied Physiology, Cells, Tissues, & Organs, Clinical Science, European Journal of Applied Physiology, Hormone Research, International Journal of Sport Nutrition & Exercise Metabolism, Journal of Applied Physiology, Journal of Athletic Training, Journal of Biomechanics, Journal of Musculoskeletal and Neuronal Interactions, Journal of Physiology, Journal of Strength and Conditioning Research, Medicine and Science in Sport and Exercise, Muscle and Nerve, PloS ONE, Research Quarterly for Exercise and Sport, and Sports Medicine.

Served on grant proposal review panels for the American Federation for Aging Research, the National Institute for Occupational Safety and Health, the United Kingdom Government/Wellcome Trust, and the U.S. Army Medical Research and Materiel Command.


Fellow, American College of Sports Medicine
Member, American Physiological Society

 Awards and Honors

  • Research publications have been cited over 6100 times in the scientific literature.
  • Recipient of three Department of Defense and three National Institutes of Health R01 research grants
  • Recipient of 2012 Henry J. Montoye Scholar Award in the Southeast Chapter of the American College of Sports Medicine